Diane Abbott has been sacked by Labour leader Ed Miliband as he continues to reshuffle his team ahead of the next election.
Ms Abbott, who ran for the party leadership against Mr Miliband in 2010, lost her job as shadow public health minister.
The move follows high profile sackings by the party leader on Monday, in what was dubbed a "cull of the Blairites" by critics.
Shadow work and pensions secretary Liam Byrne and shadow education secretary Stephen Twigg both lost their frontbench jobs.
Ms Abbott reportedly said "I'll live, these things happen" after being told she was out, adding: "I think Ed wanted more message discipline."
She was the first black woman MP when she joined Parliament in 1987 and is considered a potential future candidate for Mayor of London.
Norman Baker: No rift with Theresa May A prominent figure on Labour's left wing, the politician is regarded as one of the most independent-minded MPs in the House of Commons.
She recently questioned the party's approach on immigration and also suggested Mr Miliband was being swayed by opinion polls.
In the summer, she caused controversy by declaring she would quit if the party backed military action in Syria in a Commons vote.
Labour ultimately decided to vote against the Government, inflicting a humiliating defeat on David Cameron, so Ms Abbott stayed put.
On the coalition side, new Home Office Minister Norman Baker has scotched reports of a clash between him and his new boss Theresa May.
The Home Secretary was said to be "spitting tacks" after Mr Baker was moved to her department to replace Jeremy Browne.
But the Lib Dem insisted they had had a "very friendly chat" following his appointment and would be able to work well together despite coming "from different places".
Mr Baker, who wrote a book linking the security services to a cover-up over the death of David Kelly, also insisted he "entirely respects" the work of British spies.
Senior Lib Dem Danny Alexander predicted Mr Baker would be an "excellent" Home Office minister and would allow the party to "sharpen our campaigning edge" in that area.
Meanwhile, Tory Robert Syms' estranged wife Fiona has attracted attention after reacting with fury to him losing his post in the Whips' Office.
She wrote on Twitter: "PM just fired father of my kids over the phone. Gave up chairmanship of a cttee to be a whip, worked hard and was widely acknowledged to be a good whip.
"He was utterly gracious and took it like a man, I am beyond furious. Loyalty counted for nothing."
She joked her children had "defected to Labour" and that in her husband's position she would be "fixing up drinks" with rebels, but said the Poole MP would remain "super loyal".
She later updated her Twitter profile to describe herself as an "ex-wife with big mouth" who was "prone to extraordinary outbursts", although this has now been removed.
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